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[표제지 등]=0,1,2

제출문=0,3,1

요약문=i,4,4

Summary=v,8,5

Contents=i,13,2

List Of Tables=iii,15,5

List Of Figures=vii,19,3

목차=i,22,2

표목차=iii,24,5

그림목차=viii,29,3

제1장 연구개발과제의 개요=1,32,1

1. 연구개발의 목적=1,32,1

2. 연구개발 내용 및 범위=1,32,1

3. 연구개발의 필요성=1,32,2

제2장 국내ㆍ외 기술개발 현황=3,34,1

1. 이온빔에 의한 돌연변이 육종=3,34,5

2. 국외 돌연변이 육종 현황=8,39,3

3. 국내 돌연변이 육종 현황=11,42,1

4. 국내ㆍ외 기술수준 비교=12,43,1

제3장 연구개발 수행 내용 및 결과=13,44,1

제1절 연구내용 및 방법=13,44,1

1. 채소작물=13,44,1

가. 공시 재료=13,44,1

나. 방사선 종류 및 처리 선량=13,44,1

다. 방사선 처리=13,44,1

라. M₁세대의 육성, 생육조사 및 화분조사=13,44,3

마. 엽록소 돌연변이=15,46,1

바. 형태적 돌연변이=15,46,2

사. 내한성 돌연변이 선발=16,47,2

아. 돌연변이 계통의 DNA 분석=17,48,2

자. 유망 돌연변이 계통의 실용화을 위한 시험교잡=18,49,1

2 . 화훼작물=18,49,1

가. 화훼식물의 수집 및 육성=19,50,1

1) 식물재료=19,50,1

2) 기내식물체배양=19,50,2

나. 이온 빔 조사=20,51,1

1) (12)C 이온빔 조사 ( 135 MeV/μ, LET 22 KeV/μm)(이미지 참조)=20,51,1

2) (20)Ne 이온빔조사(이미지 참조)=20,51,2

다. 기호성 화훼류 변이체선발과 특성조사=21,52,1

라. 이온빔 조사에 따른 화훼작물의 유전자 변이체 해석=21,52,1

1) DNA 추출=21,52,1

2) RAPD=21,52,1

3) AFLP=21,52,2

4) 유전자 결손 양식 검토=22,53,1

제2절 연구결과 및 고찰=23,54,1

1. 채소작물=23,54,1

가. 배추 (Brassica Camperstris)=23,54,1

1) 방사선이 M1세대의 생육에 미치는 영향=23,54,15

2) 엽록소 돌연변이=38,69,5

3) 배추의 형태적 돌연변이=42,73,6

4) 배추의 내한성 변이체 선발=47,78,3

5)/4) 배추 돌연변이 계통의 DNA 분석=49,80,3

6)/5) 배추 유망 선발계통의 실용화를 위한 F1시험교배 검정=51,82,4

나. 무(Raphanus sativus)=54,85,1

1) M1세대의 생육에 미치는 영향=54,85,8

2) 무 엽록소 돌연변이=61,92,4

3) 무 형태적 돌연변이=65,96,4

4) 무 내한성돌연변이=69,100,3

5) 무 돌연변이 계통의 DNA 분석=71,102,3

6) 무 유망선발계통의 품종화를 위한 교배조합=73,104,2

다. 고추(Capsicum Annum)=75,106,1

1) 이온 빔이 고추의 M1세대 생육에 미치는 영향=75,106,8

2) 이온 빔 처리가 고추의 돌연변이 유기에 미치는 효과=82,113,7

2. 화훼작물=89,120,1

가. 피조사체 및 조사후 화훼식물의 육성=89,120,3

나. 이온빔조사에 따른 식물체의 생장특성=91,122,9

다. 변이체 선발=100,131,2

라. 변이체집단해석=102,133,1

1) RAPD분석과 유전자산물의 클로닝=102,133,1

2) AFLP 분석=102,133,2

(1) RAPD분석=104,135,2

(2) AFLP 분석=105,136,2

3) 배수성조사=106,137,3

제3절 종합고찰=108,139,9

제4장 연구개발 목표 달성도 및 관련분야 기여도=117,148,1

제1절 연구목표 및 연구개발목표의 달성도=117,148,1

제2절 관련분야 기술 발전등에의 기여도 및 파급효과=118,149,1

제5장 연구개발결과의 활용계획=119,150,1

제6장 참고문헌=120,151,6

서지정보양식=126,157,2

표목차

Table 1. 각국별 돌연변이 유기에 의한 육성된 품종 수(Fao/Iaea, 2002)=10,41,1

Table 2. 작물 유형별 전세계의 돌연변이 유기 육성 품종수(Fao/Iaea, 2002)=10,41,1

Table 3. 우리나라 돌연변이육성품종(26품종)=11,42,1

Table 4. 공주지역 최고, 최저, 평균 기온 (2004. 2-2005.2)=16,47,1

Table 5. Effect Of Rariation On Germination Of Brassica Campestris(B1)=26,57,1

Table 6. Effect Of Rariation On Germination Of Brassica Campestris(B2)=26,57,1

Table 7. Effect Of Rariation On Germination Of Brassica Campestris(B3)=26,57,1

Table 8. Effect Of Rariation On Germination Of Brassica Campestris(B4)=27,58,1

Table 9. Effect Of Rariation On Germination Of Brassica Campestris(B5)=27,58,1

Table 10. Effect Of Rariation On Germination Of Brassica Campestris(B6)=27,58,1

Table 11. Effect Of Rariation(Proton) On Germination Of Brassica Campestris(B7)=28,59,1

Table 12. Effect Of Rariation(Neutron) On Germination Of Brassica Campestris(B8)=28,59,1

Table 13. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B1)=29,60,1

Table 14. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B2)=29,60,1

Table 15. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B3)=29,60,1

Table 16. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B4)=30,61,1

Table 17. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B5)=30,61,1

Table 18. Effect Of Radiation On The Seeding Growth Of Brassica Campestris(B6)=30,61,1

Table 19. Effect Of Rariation(Proton) On The Seeding Growth Of Brassica Campestris(B7)=31,62,1

Table 20. Effect Of Rariation(Neutron) On The Seeding Growth Of Brassica Campestris(B8)=31,62,1

Table 21. Flowering Date Of Brassica Campestris Irradiated With Ion Beams And Gamma Ray=32,63,1

Table 22. Pollen Fertilities(%) Of Brassica Campestris Irradiated With Ion Beams And Gamma Ray=32,63,1

Table 23. Chlorophyll Mutation Line And Rate Of M₂Generation Of Brassica Campestris Irradiated With Ion Beam And Gamma Lay=41,72,1

Table 24. Chlorophyll Mutation Plant And Rate Of M₂Generation Of Brassica Campestris Irradiated With Ion Beam And Gamma Lay=41,72,1

Table 25. Chlorophyll Mutation Frequency Of Brassica Campestris Treated With Gamma Lay=42,73,1

Table 26. Variation Mutation Line And Rate Of M₂ Generation Of Brassica Campestris Irradiated With (12)C Ion Beam And Gamma Ray(이미지 참조)=45,76,1

Table 27. Variation Mutation Line And Rate Of M₂ Generation Of Brassica Campestris Irradiated With Ion Beam And Gamma Ray=45,76,1

Table 28. Variation Mutation Line And Rate Of M₂ Generation Of Brassica Campestris Irradiated With Ion Beam And Gamma Ray=46,77,1

Table 29. Variation Mutation Line And Rate Of M₂ Generation Of Brassica Campestris Irradiated With Ion Beam And Gamma Ray=46,77,1

Table 30. Frequency Of Variation In M₂ Generation Of Brassica Rapa Var Pekinensis Irradiated With (12)C Ion Beam(이미지 참조)=47,78,1

Table 31. Mutant Plants Of Cold Hardiness And Rate Of M₂ Generation Of Brassica Rapa Ivar Pekinensis Irradiated With Gamma Ray And Ion Beam=49,80,1

Table 32. Comparison Of Band Numbers Between Control (B1) And Their Mutants=51,82,1

Table 33. 배추 유망 돌연변이 계통의 F1 교배조합=54,85,1

Table 34. Effect Of Radiation On Generation Of Raphanus Sativus(R1)=55,86,1

Table 35. Effect Of Radiation On Generation Of Raphanus Sativus(R2)=56,87,1

Table 36. Effect Of Radiation On Generation Of Raphanus Sativus(R3)=56,87,1

Table 37. Effect Of Radiation On Generation Of Raphanus Sativus(R4)=56,87,1

Table 38. Effect Of Radiation On Generation Of Raphanus Sativus(R5)=57,88,1

Table 39. Effect Of Radiation On The Seeding Growth Of Raphanus Sativus(R1)=57,88,1

Table 40. Effect Of Radiation On The Seeding Growth Of Raphanus Sativus(R2)=57,88,1

Table 41. Effect Of Radiation On The Seeding Growth Of Raphanus Sativus(R3)=58,89,1

Table 42. Effect Of Radiation On The Seeding Growth Of Raphanus Sativus(R4)=58,89,1

Table 43. Effect Of Radiation On The Seeding Growth Of Raphanus Sativus(R5)=58,89,1

Table 44. Pollen Fertilities(%) Of Raphanus Sativus Irradiated With Ion Beams And Gamma Ray=59,90,1

Table 45. Chlorophyll Mutation Line And Rate Of M₂Generation Of Raphanus Sativus Irradiated With Ion Beam And Gamma Lay=63,94,1

Table 46. Chlorophyll Mutation Line And Rate Of M₂Generation Of Raphanus Sativus Irradiated With Ion Beam And Gamma Lay=64,95,1

Table 47. Chlorophyll Mutation Frequency Of Raphanus Sativus Treated With Gamma Lay=64,95,1

Table 48. Variation Mutation Line And Rate Of M₂ Generation Of Raphanus Sativus Irradiated With Gamma Ray=66,97,1

Table 49. Variation Mutation Line And Rate Of M₂ Generation Of Raphanus Sativus Irradiated With Gamma Ray=67,98,1

Table 50. Variation Mutation Line And Rate Of M₂ Generation Of Raphanus Sativus Irradiated With Gamma Ray=68,99,1

Table 51. Variation Mutation Line And Rate Of M₂ Generation Of Raphanus Sativus Irradiated With Gamma Ray=68,99,1

Table 52. Mutant Plants Of Cold Hardiness And Rate Of M₂ Generation Of Raphanus Sativus Irradiated With Gamma Ray=70,101,1

Table 53. Frequency Of Variation In M₂ Generation Of Raphanus Sativus Irradiated With (12)C Ion Beam(이미지 참조)=71,102,1

Table 54. 무 돌연변이 계통의 실용화를 위한 교배조합=74,105,1

Table 55. Effect Of Radiation On The Germination Of Capsicum Annuum(Pe1) Irradiated With Gamma Ray=78,109,1

Table 56. Effect Of Radiation On The Germination Of Capsicum Annuum(Pe2) Irradiated With Gamma Ray=79,110,1

Table 57. Effect Of Radiation On The Seeding Growth Of Capsicum Annuum(Pe1) Irradiated With Gamma Ray=79,110,1

Table 58. Effect Of Radiation On The Seeding Growth Of Capsicum Annuum(Pe2) Irradiated With Gamma Ray=79,110,1

Table 59. Pollen Fertilities(%) Of Capasicum Annum Irradiated With Ion Beams And Gamma Ray=82,113,1

Table 60. Chlorophyll Mutation Frequency Of Capsicum Annum (Pe3) Treated With (12)C Ion Beam(이미지 참조)=86,117,1

Table 61. Line Of M₂Generation Of Capasicum Annuum(Pe1) Irradiated With Gamma Ray=87,118,1

Table 62. Line Of M₂Generation Of Capasicum Annuum(Pe2) Irradiated With Ion Beam And Gamma Ray=87,118,1

Table 63. Frequency Of Mutation Of Capasicum Annuum(Pe2) Irradiated With (12)C Ion Beam And Gamma Ray In M₂ Generation(이미지 참조)=88,119,1

Table 64. Frequency Of Mutation Of Capasicum Annuum(Pe2) Irradiated With (12)C Ion Beam And Gamma Ray In M₂ Generation(이미지 참조)=88,119,1

Table 65. Effect Of Sugar Concentration On Growth Characters Of Bulb In Lilium Tsintauense Var. Flavum=89,120,1

Table 66. Growth Pattern Of Lilium Tsintauense Var. Flavum After Proton Beam Irradiation In 8 Weeks Culture=91,122,1

Table 67. Growth Pattern Of Lilium Tsintauense Var. Flavum After Proton Beam Irradiation In 90 Days Culture=92,123,1

Table 68. Effect Of X-Ray Irradiation On Height And Shoot Number Of Lilium Tsintauense Var. Flavum Cultured In Vitro=92,123,1

Table 69. Growth Pattern Of Lilium Longiflorum After Proton Beam Irradiation In 90 Days Culture=93,124,1

Table 70. Effect Of Irradiation X-Ray On Height And Shoot Number Of In Vitro Culture Lilium Longiflorum=94,125,1

Table 71. Growth Pattern Of Rose After Proton Beam Irradiation In 4 Weeks Culture=95,126,1

Table 72. Effect Of Irradiation X-Ray On Height And Shoot Number Of In Vitro Culture Rose=95,126,1

Table 73. Growth Pattern Of Phalaenopsis After Proton Beam Irradiation In 12 Weeks Culture=96,127,1

Table 74. Growth Pattern Of Phalaenopsis After Proton Beam Irradiation In 12 Weeks Culture=96,127,1

Table 75. Growth Pattern Of Chrysanthemum Cutting After Proton Beam Irradiation In 4 Weeks Culture=98,129,1

Table 76. Shoot Induction From Leaf Segment Of Proton Beam Irradiated Chrysanthemum=98,129,1

Table 77. Polymorphism By Aflp Analysis Among The In Vitro Culture Bulblets By Lilium Tsintauense Var. Flavum=103,134,1

Table 78. Polymorphism By Sucrose Concentration Among The In Vitro Culture Bulblets By Lilium Tsintauense Var. Flavum=103,134,1

Table 79. Neucleotide Sequence And Of Random Primer(Operon Technologies Inc.) And Amplified Dna Bands=104,135,1

Table 80. Number Of Total Bands Observed In Orchid Rhizomes By Aflp Analysis=106,137,1

Table 81. Polyploidylevel Of Proton Beam Irradiated Chrysanthemum=108,139,1

그림목차

Figure 1. Transplanting And Bagging For Seef Setting In Vinyl House=14,45,1

Figure 2. Temperature Of Kong-Ju Area From November To December In 2003(Agricultural Technology And Extension Center)=17,48,1

Figure 3. Effect Of Radiation On The Seeding Growth Of Brassica Campestries=25,56,1

Figure 4. Pollen Ferilities Of Brassica Campestris(B1)×400 (A: Control, B: 500Gy, C:750Gy, D: 1000Gy)=33,64,1

Figure 5. Pollen Ferilities Of Brassica Campestris(B3)×400 (A: Control, B: 500Gy, C:750Gy, D: 1000Gy)=34,65,1

Figure 6. Pollen Ferilities Of Brassica Campestris(B4)×400 (A: Control, B: 500Gy, C:750Gy, D: 1000Gy)=35,66,1

Figure 7. Pollen Ferilities Of Brassica Campestris(B6)×400=36,67,1

Figure 8. Pollen Ferilities Of Brassica Campestris(B3)×400=37,68,1

Figure 9. Chlorophyll Mutantion Of Chinese Cabbage Irradiated With Gamma Ray And (12)C Ion Beam(이미지 참조)=39,70,1

Figure 10. Homozygous And Heterozygous Chlorophyll Mutant In M₃ Generation Of Brassica Rapa Var Pekinensis Irradiated With (12)C Ion Beam And Gamma Ray(이미지 참조)=40,71,1

Figure 11. Various Morphological Mutants Of Chinese Cabbage Irradiated With Gamma Ray And (12)C Ion Beam(이미지 참조)=43,74,1

Figure 12. Homozygous Plant Tpe Mutant Of Brassica Rapa Var Pekinensis Irradiated With (12)C Ion Beam And Gamma Ray In M₃ Generation(이미지 참조)=44,75,1

Figure 13. Various Of Cold Hardiness Selected In M₂ Generation Of Brassica Rapa Var Pekinensis Irradiated With (12)C Ion Beam(이미지 참조)=48,79,1

Figure 14. Rapd Profiles Of Brassica Campestris With Operon=50,81,1

Figure 15. Various Promising Mutants For Commercial Varieties Of Brassica Rapa Var Pekinensis In M₃ Generation=52,83,1

Figure 16. Diagram Of Test For Crossing Ability Of Mutants For Commercial Varieties=53,84,1

Figure 17. Effect Of Radiation On The Root Growth Of Raphanus Sativus=55,86,1

Figure 18. Pollen Fertilities Of Raphanus Sativus(R1)×400=59,90,1

Figure 19. Pollen Fertilities Of Raphanus Sativus(R2)×400=60,91,1

Figure 20. Pollen Fertilities Of Raphanus Sativus(R3)×400=60,91,1

Figure 21. Pollen Fertilities Of Raphanus Sativus(R5)×400=61,92,1

Figure 22. Chlorophyll Mutantion Of Radish Irradiated With Gamma Ray=62,93,1

Figure 23. Various Plant Type Mutation(A - F) Of Radish Irradiated With Gamma Ray=65,96,1

Figure 24. Radish Mutant Of Cold Hardiness Selected In M2 Generation On 29Th December In 2003=69,100,1

Figure 25. Rapd Profiles Of Brassica Campestris With Operon=72,103,1

Figure 26. Various Leaf And Root Type Mutants Of Radish In M₃ Generation=73,104,1

Figure 27. Effect Of Radiation On The Seeding Growth Of Campestries Annum=77,108,1

Figure 28. Pollen Fertilities Of Capasicum Annuum(Pe1)×400(A: Control, B: 250Gy, C:500Gy)=80,111,1

Figure 29. Pollen Fertilities Of Capasicum Annuum(Pe1)×400=81,112,1

Figure 30. Fruit Type Mutation Of Capcicum Annum Irradiated With (12)C Ion Beam And Gamma Ray(이미지 참조)=85,116,1

Figure 31. Fruit Type Mutation Of Capcicum Annum Irradiated With (12)C Ion Beam And Gamma Ray(이미지 참조)=86,117,1

Figure 32. Profiles For Multiple Propagation Of Lilium Tsintauense Var. Flavum Cultured On Ms Media Containing 1.0Mg/L Ba, 0.1Mg/L Naa=90,121,1

Figure 33. Profiles Of Lilium Longiflorum Culture On Pot=90,121,1

Figure 34. Profiles Of Shoot Growth Of Proton Beam Irradiated Lilium Longiflorum=93,124,1

Figure 35. Profiles Of Shoot Growth Of Proton Beam Irradiated Phalaenopsis. Upper: 500Gy To 300Gy Irradiation, Lower: 50Gy To 100Gy Irradiation=97,128,1

Figure 36. Profiles Of Shoot Growth Of Proton Beam Irradiated C,Goeringii=97,128,1

Figure 37. Profiles Of Proton Beam Irradiated Plants (Upper: Pm, Middle: Zw , Lower: Uy). These Shoe Difference Of Proton Susceptibility By Lines In The Case Of Cutting Culture=99,130,1

Figure 38. Profiles Of Shooting Of Proton Beam Irradiated D. Grandiflorum=99,130,1

Figure 39. Profiles Of Variegated Bulb And Leaf Of Lily Induced By Proton Beam Irradiation, Left: Variegated Plants. Right: In Vitro Propagated Bulblets=100,131,1

Figure 40. Profiles Of Flower Color Changed Rose Induced=100,131,1

Figure 41. Profiles Of Rhizomes Induction Of Orchids 18 Months After C Ion-Beam Irradiation=101,132,1

Figure 42. Profiles Of Variegated Leaves Of Chrysanthemum Derived From 20Gy Of Proton Beam Irradiated Cuttings=101,132,1

Figure 43. Profiles Of Rapd Products And Cloned Dna Fragments=102,133,1

Figure 44. Rapd Profiles Of Orchid Rhizomes After (12)C Beam Irradiation(이미지 참조)=105,136,1

Figure 45. Aflp Patterns Of Orchid Rhizome With The Primers=105,136,1

Figure 46. Aflp Patterns Of Orchid Rhizome With The Primers=107,138,1

Figure 47. Profile Of Polyploidy Of Chrysanthemum=107,138,1

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